To the taxonomy of Agapanthia dahlii (Richter, 1821) (Coleoptera, Cerambycidae) subspecies distributed from West Europa to Russia, Near East and Central Asia with several new descriptions Текст научной статьи по специальности «Биологические науки»

http://zoobank.org/urn:lsid:zoobank.org:pub:4DAE5E92-C9ED-4FBA-AFE2-007D9E789503

DOI: 10.24412/2226-0773-2025-14-1-29-79

EDN: QSBOZA

To the taxonomy of Agapanthia dahlii (Richter, 1821) (Coleoptera, Cerambycidae) subspecies distributed from West Europa to Russia, Near East and Central Asia with several new descriptions

M.A. Lazarev

Free Economic Society of Russia, Department of Scientifics Conferences and

All-Russian Projects

Tverskaya str., 22b, Moscow 125009 Russia

e-mail: [email protected]; [email protected]

ORCID 0000-0002-4040-0987

Key words: Coleoptera, Cerambycidae, taxonomy, zoogeography, new subspecies, new synonyms, subspecies key.

Abstract. Eight new subspecies are described: Agapanthia (Epoptes) dahlii chemalensis ssp. n. (type locality: Chemal, Republic of Altay, Russia), A. (E.) d. setosa ssp. n. (type locality: Eğridir environs, Isparta, Turkey), A. (E.) d. grossicornis ssp. n. (type locality: Çorum, Turkey), A. (E.) d. zaysanensis ssp. n (type locality: Zaysan Lake environs), A. (E.) d. zhidkovi ssp. n. (type locality: 55 km S Makanchi, Kazakhstan), A. (E.) d. vishnyakovi ssp. n. (type locality: Kitab Natural reserve, Zerafshan mountain ridge, Kashkadarya Region, Uzbekistan), A. (E.) d. krivosheinae ssp. n. (type locality: Yangier, Syrdarya Region, Uzbekistan), A. (E.) d. revadensis ssp. n. (type locality: Revad environs, Zerafshan valley, Tadzhikistan). Two synonyms are proposed: A. (E.) pustulifera Pic, 1905 = A. (E.) mutinensium Sama & Rapuzzi, 2010, syn. nov. = A. (E.) subsimplicicornis Sama & Rapuzzi, 2010, syn. nov. New distributional data are represented for 3 subspecies. A key for all subspecies is proposed.

Introduction.

Agapanthia dahlii (Richter, 1821) is a very complicated taxon of many rather different subspecies. According to the last Cerambycidae Palaearctic Catalogue (Danilevsky, 2020), it included 19 subspecies. Recently one new subspecies was described (A. d. efimovi Danilevsky, 2021) and the names of 7 species were downgraded to subspecies rank (Lazarev, 2024). Now the species consists of 33 subspecies. Below 8 new subspecies are described and 2 names are accepted as new synonyms.

Materials and methods

Material was collected manually. Specimens used in morphological studies were killed by ethyl acetate. All photographs were taken with Canon PowerShot G10 digital camera equipped with Cannon Zoom lens 5X IS 6.1-30.5 mm 1:2.8-4.5 and microscope AmScope SM745NTP. The illustrations were edited with Adobe Photoshop 7.0 and Helicon Focus 3.20.

Acronyms of collections:

DE - collection of Dmitry A. Efimov (Kemerovo, Russia);

DN - collection of David Navrátil (Litomyšl, Czech Republic);

KH - collection of Karel Hodek (Brno, Czech Republic);

MD - collection of Mikhail L. Danilevsky (Moscow, Russia);

MH - collection of Michal Holomčík (Lužice, Czech Republic);

ML - collection of Maxim A. Lazarev (Moscow, Russia);

SM - collection of Sergey V. Murzin (Moscow, Russia);

VU - collection of Vadim E. Ustinov (Moscow, Russia);

ZIN - collection of Zoological Institute of the Russian Academy of Sciences (Saint-Petersburg, Russia);

ZMM - collection of Zoological Museum of Moscow University (Russia).

Results

A list of Agapanthia (Epoptes) dahlii (Richter, 1821)

groups of subspecies

The subspecies could be joined in several related groups on the bases of geographical distribution and natural connections. Subspecies of the first group “A. d. dahlii” are distributed from Europe to Siberia and usually have densely pubescent elytra and setae tufts of 3rd antennal joints. Subspecies of the second group “A. d. walteri” are distributed in Transcaucasia and neighbor regions and usually characterized by spotted elytra and well developed antennal setae tufts. Subspecies of the third group “A. d. sicula” are distributed in West Europe and Near East and usually have rather glabrous elytra. Subspecies of the fourth group “A. d. muellneri” are distributed in Central Asia and are characterized by well-developed elytral pubescence.

The records of A. dahlii from China and Mongolia could be connected with unknown subspecies not described yet, or to wrong identifications. No A. dahlii from China and Mongolia are known to the author.

I. group A. d. dahlii (Richter, 1820)

1. A. d. dahlii (Richter, 1821) -Map 1: No 1.

Type locality. “Hungaria” - after original description.

2. A. d. kuleshovi Danilevsky, 2018 - Map 1: No 2.

Type locality. Russia, Tomsk Region, Belousovo environs, 56°18’13"N, 85°11’53"E.

3. A. d. efimovi Danilevsky, 2021 - Map 1: No 3.

Type locality. Russia, Kemerovo Region, Prokopyevsk District, Karakan Mt. Ridge., 6-7 km from Tykhta.

4. A. d. chemalensis ssp. n. - Map 1: No 4.

Type locality. Russia, Republic of Altay, Chemal.

5. A. d. calculensis Lazarev, 2013 - Map 1: No 5.

Type locality. North-east Kazakhstan, the valley of the Sibinka River about 40 km south of Ust-Kamenogorsk, 49°40'27.56"N, 82°39'13.12"E.

II. group A. d. walteri Reitter, 1898

6. A. d. walteri Reitter, 1898 - Map 1: No 6.

Type locality. Turkey, Erzurum.

7. A. d. nitidipennis Holzschuh, 1984 - Map 1: No 7.

Type locality. Georgia, environs of the Jvari monastery (41°50′19″N, 44°44′02″E) north of Tbilisi.

8. A. d. rubenyani Lazarev, 2013 - Map 1: No 8.

Type locality. South Armenia, Megri District, mountains above Shvanidzor, 39°13'N, 46°22'44''E, 1600 m.

9. A. d. ismailovae Lazarev, 2013 - Map 1: No 9.

Type locality. Russia, North Cucasus, Dagestan, Rutul environs (41°32'N, 47°25'E).

10. A. d. persica Semenov, 1893 - Map 1: No 10.

Type locality. Iran, Eastern and Central Mazanderan province.

11. A. d. lenkorana Lazarev, Plewa & Jaworski, 2016 - Map 1: No 11.

Type locality. Azerbaijan, Jalal-Abad District, Andreevka environs (Karazenjir, 39°16'N, 48°30'E, 10 m).

12. A. d. golestanica Lazarev, Plewa & Jaworski, 2016 - Map 1: No 12.

Type locality. Iran, Golestan province, 60 km east Minudasht, 460 m, 37°21'36"N 55°55'48"E.

13. A. d. salviae Holzschuh, 1975 - Map 1: No 13.

Type locality. Iran, Elburz, south side, 10 km north of Karaj.

14. A. d. transcaspica Pic, 1900 - Map 1: No 14.

Type locality. Turkmenistan, Ashgabat.

III. group A. d. sicula Ganglbauer, 1884

15. A. d. sicula Ganglbauer, 1884 - Map 1: No 15.

Type locality. Italy, Sicilia.

16. A. d. schurmanni Sama, 1979 - Map 1: No 16.

Type locality. Greece, Kastoria.

17. A. d. malmerendii Sama, 1981 - Map 1: No 17.

Type locality. Italia, Romagna, Portico di Romagna [44°1'N, 11°46'E].

18. A. d. lateralis Ganglbauer, 1884 - Map 1: No 18.

Type locality. Turkey, Istanbul Province, Istanbul (Constantinopel).

19. A. d. pustulifera Pic, 1905 - Map 1: No 19.

Type locality. Israel, Jerusalem.

20. A. d. kindermanni Pic, 1905 - Map 1: No 20.

Type locality. Southern Turkey.

21. A. d. setosa ssp. n. - Map 1: No 21.

Type locality. Turkey. Isparta, Eğridir environs.

22. A. d. grossicornis ssp. n. - Map 1: No 22.

Type locality. Turkey, Çorum.

IV. group A. d. muellneri Reitter, 1898

23. A. d. iliensis Danilevsky, 2018 - Map 1: No 23.

Type locality. Kazakhstan, Almaty Region, southern environs of Lake Sorbulak (43°33'57.65"N, 76°36'24.93"E), 670 m.

24. A. d. alexandris Pic, 1901 - Map 1: No 24.

Type locality. Kyrgyzstan, Kyrgyz Ridge (formerly Alexander Ridge); the type most likely comes from the western (Kazakh) part of the ridge.

25. A. d. muellneri Reitter, 1898 - Map 1: No 25.

Type locality. Uzbekistan, Tashkent.

26. A. d. alaiensis Kratochvíl, 1985 - Map 1: No 26.

Type locality. Southern Kyrgyzstan at the southern border of the Fergana Valley, Kadamjai (40°7'44"N, 71°43'26"E).

27. A. d. zaysanensis ssp. n. - Map 1: No 27.

Type locality. East Kazakhstan, Zaysan lake environs, Zhemeney River.

28. A. d. zhidkovi ssp. n. - Map 1: No 28.

Type locality. Kazakhstan, eastern shore of Alakol lake, 60 km S Makanchi, 408 m, 46°15'24.54"N, 82°12'55.68"E.

29. A. d. lepsyensis Danilevsky, 2018 - Map 1: No 29.

Type locality. Kazakhstan, Lepsy river, 7 km northeast Koilyk (formerly Antonovka), 45°41'36.22"N, 80°17'58.94"E.

30. A. d. vishnyakovi ssp. n. - Map 1: No 30.

Type locality. Uzbekistan, Kashkadarya Region, Zerafshan mountain ridge, Kitab Natural reserve (about 39°10'30"N, 67°18'43"E).

31. A. d. krivosheinae ssp. n. - Map 1: No 31.

Type locality. Uzbekistan, Syrdarya Region, Yangier.

32. A. d. revadensis ssp. n. - Map 1: No 32.

Type locality. Tadzhikistan, Zerafshan valley, Revad environs.

33. A. d. ustinovi Danilevsky, 2013 - Map 1: No 33.

Type locality. Tadzhikistan, Pamir, Poshkharv environs [38°24'1"N, 71°9'18"E].

Agapanthia (Epoptes) dahlii (Richter, 1821)

Saperda cardui, Fabricius, 1775: 186 - “Habitat in Europae australioris carduis”; 1801: 325 (misapplied) - “Habitat in Europae australioris carduis”; Herbst, 1784: 94; Olivier, 1795: N68, p. 9 - “dans les départemens meridionaux de la France”, “l’Allemagne”.

Saperda nigricornis Fabricius, 1793: 314 (preoccupied) - “Habitat in Europae australioris”.

Saperda dahlii Richter, 1821: pl. 12 - “Hungaria”.

Agapanthia cardui, Mulsant, 1839: 175 - “la France méridionale et témperée”

Agapanthia lineatocollis, Mulsant, 1863: 358 (misapplied) - France, “les zones témperées ou méridionales”.

Agapanthia gyllenhali Ganglbauer, 1883: 190 (= cardui, F. = lineatocollis, Muls.) - “E. md.”.

Agapanthia dahli, Ganglbauer, 1884: 541 - “Mittel- und Süd-Europa, Caucasus, Kleinasien, Syrien”; Reitter, 1898: 133 - “Mittel- und Südeuropa, Kaukasus; dann angeblich auch in Kleinasien und Syrien”; 1913: 66 - “Süddeutschl., Nassau, Böhmen”; Stierlin, 1898: 498 - “Genf, Aarau, Dübendorf, Glarner Alpen”; Everts, 1901: 386; Csiki, 1905: 64 - Hungary; Schaufuß, 1916: 876 - “Mittel- u. Südeuropa”; Plavilstshikov, 1927: 61 - “au Caucase, en Transcaucasie, dans le Turkestan boréal, la Sibérie mér. occ., et la Russie d’Europe moyenne et méridionale”; 1929: 103, part. (= kindermanni Pic, 1905); 1932: 194; 1965: 416 - the south and center of the European part of the USSR, Caucasus, Western Siberia; Chernyshev, 1930: 12 - Bryansk province; Esterberg, 1935: 197 - Gorky and Kirov regions; Zaitzev, 1954: 18, part. - Georgia: (Borjomi, Eldar, Tbilisi, Manglisi, Gagra), All Transcaucasia, Europe, Siberia, Western. Asia; Yablokov-Khnzoryan, 1961: 78 - Europe, including Armenia; Breuning, 1961: 185, part. - “Eur. centr. et mer., As. occ. et centr.”; Abai, 1969: 53, part. - Iran: Gorgan, Khorassan; Kostin, 1973: 224 - Kazakhstan; Shernin, 1974: 181 - Urzhum, Kirov region; Villiers, 1978: 431, 434 - “Europe centrale et méridionale, Sibérie occidentale, Caucase, Proche et Moyen-Orient; Sama, 1979: 506, 511; 2003: 93 - Europe, Western Türkiye, Caucasus, Transcaucasia, Siberia to Lake Baikal; Miroshnikov, 1984: 280 (larva); Tsherepanov, 1985: 246 - Europe, Western Siberia; Novozhenov, 1987: 45 - Ilmensky Reserve; Rabil, 1992: 148 - “Forêt de la Grésigne (Tarn)”; Bense, 1995: 400-401 - Western Europe; Carrière, 1996a: 562 - “Hérault”; 1996b, 110, 111 - “Portiragnes; Pech Blanc, Etang de Vendres”; Alexandrovich et al., 1996: 48 - southeast of Belarus; Althoff & Danilevsky, 1997: 40 - Europe; Kasatkin & Arzanov, 1997: 67 - Rostov Region, Volgograd Region, Astrakhan Region, Kalmykiya, Krasnodar Region, Karachay-Cherkessia, Checheno-Ingushetia, Dagestan; Matveev, 1998: 87 - Mari El, Tatarstan, Kirov, Nizhny Novgorod Regions; Secchi, 1998: 227 - “Loiret: Meung-sur-Loire”; Kovács, 1998: 254 - Hungary; Efimov, 2001: 69 - Kemerovo Region; Hua, 2002: 191, part. - China; Former USSR, Mongolia; Brustel et al., 2003: 452; Isaev et al.., 2004: 41 - Tatarstan, Ulyanovsk and Samara Regions; Diego Barquín & Martínez-Porres Cáceres, 2005: 145 - Spain: Cantabria; Sautière, 2005: 21 - Montlouis-sur-Loire (Indre-et-Loire); Verdugo, 2008: 484 - Andalucía: Calar Alto, término municipal de Bacares, Almería; Mouthiez & Péru, 2008: 110 - Loiret; Sama et al., 2008: 122, part. - “absent in Iran”; Kadyrbekov & Tleppaeva, 2008: 54 - tugai forests of Semirechye and the steppe belt of the Dzhungar Alatau and Northern Tien Shan; Tiberghien, 2010: 63 - Zaragoza; Gnjatović & Žikić, 2011: 36 - Montenegro; Hernández, 2011: 257 - Lerma (Burgos); Danilevsky, 2012: 153 - Palaearctic, including Belgium but excluding Korea; Zamoroka et al., 2012: 1167 - Western Podillya, Ukraine (East Pokuttya, Khotyn Eminence); Švácha & Lawrence, 2014: figs 2.4.20 T (larva), 2.4.32 M (nymph female); Dobrosavljević & Mihajlović, 2014: 26 - Serbia; Pavićević et al., 2015: 83 - Serbia; Kulenko, 2015: 1104 - Russia, Samara Region: (Tolyatti, Zhigulevsk); Molnar et al., 2016: 49 - Hungary (Fundoklia Valley); Danilevsky, 2017: 28 - South-West Siberia; Touroult et al., 2019: 107 – France; Nikitsky, 2019: 575 - Moscow Region.

Agapanthia dahlii, Seidlitz, 1891b.: 850 (= cardui, F. = lineatocollis, Muls.) - “In Eur. bis Ostpr. (?)”.

Agapanthia (s. str.) dahli, Aurivillius, 1923: 461 - Mittel- und Süd-Europa; Martynov & Pisarenko, 2004: 64 - Lugansk and Donetsk Regions.

Agapanthia (Agapanthiella) dahlii, Pesarini & Sabbadini, 2004: 127, part.; Danilevsky, 2006: 49 - Moscow Region; Özdikmen, 2007: 348, 392 - Europe, European Russia, European Kazakhstan, Siberia, Central Asia, ?Mongolia, China, Caucasus, Transcaucasia, Near East, Turkey, Iran; Listvyagova et al., 2013: 28 - Republic of Khakassia, Krasnoyarsk Krai.

Agapanthia (s. str.) dahli dahli, Bartenev, 2004: 387 - Europe, Caucasus and Transcaucasia, “Northern Iran, Syria, Turkey, Palestine, Northwestern Kazakhstan, Uzbekistan, Turkmenistan, Southwestern Siberia.

Agapanthia (Epoptes) dahli, Löbl & Smetana, 2010: 215; Drumont & Leduc, 2011: 293, 295 - “Belgique: province de Namur”; Shapovalov, 2012: 184 - Russia east to approximately Lake Baikal, Europe, “Caucasus, Central Asia, possibly also northwestern Mongolia and northwestern China”; Steiner & Schmid, 2013: 2 - “Griechenland”; Klausnitzer et al., 2016: 557 - Mitteleuropa; Stolbov et al., 2019: 209 - Russia (Tyumenskaya Oblast); Özdikmen, 2021: 1352; Zamoroka, 2022: 64 - Ukraine; Trócoli et al., 2023: 245 - “España (Barcelona, Catalunya): Moianès”; Danilevsky, 2020: 301, 302 - center and south of European Russia, Western Siberia, Kazakhstan, Belarus, Moldova, Ukraine, Georgia, Armenia, Azerbaijan, Turkmenistan, Uzbekistan, Tajikistan, Kyrgyzstan, Afghanistan, Iran, Turkey, China, Western Europe.

Agapanthia dahli dahli, Sláma, 1998: 350 - Czech Republic, Slovakia; Bartenev & Terekhova, 2011: 139 - Left Bank Ukraine and Crimea; Lazarev, 2013a: 443 (new Asian subspecies); 2013b: 128, 129, figs 4-5 (Samara) - Central Russia.

Agapanthia (Epoptes) dahli dahli, Mikhailov, 1999: 230 - Arkim; Georgiev, Gjonov & Sakalian, 2015: 82 - “Strandzha Mountain (Turkey: Kömürköy Köyü)”; Miroshnikov, 2011: 262; Danilevsky, 2014: 219; Gradinarov & Petrova, 2019: 70 - Bulgaria: Vrachanski Balkan Nature Park; 2020: 174 - Bulgaria: Sarnena Sredna Gora Mountains; Gradinarov et al., 2020: 106 - Bulgaria; Danilevsky, 2020: 302 - center and south of European Russia, Western Siberia, Kazakhstan, Belarus, Moldova, Ukraine, Georgia, Armenia, Azerbaijan, Turkmenistan, Uzbekistan, Tajikistan, Kyrgyzstan, Afghanistan, Iran, Turkey, China, Western Europe; Danilevsky, 2024: 153.

Agapanthia (Agapanthiella) dahli dahli, Shapovalov et al., 2006: 107 - Orenburg Region.

Agapanthia (Epoptes) dahlii, Lin & Tavakilian, 2019: 226 - “China, Mongolia, Russia, Uzbekistan, Kazakhstan, Turkey, Georgia; Europe”.

Type locality. “Hungaria” - according to the original description.

Extremely variable species with huge area. Many local forms were originally described as species or accepted as species after original publication. All forms are characterized by red basal parts of 3rd - 12th joints; pubescence of the light parts of antennal joints usually white, but sometimes yellowish; antennal length usually variable in each population from about 2 times longer than body in males protruding beyond elytral apices with 7th joint, in females - with 8th joint; elytral pubescence from pale-yellow, to dark-yellow or orange; always more or less spotted; spots can be rather contrast or partly conjugated; very rare (A. d. efimovi) elytral pubescence about uniform without spots; erect setae are usually distinct along anterior elytral half or anterior two-thirds, or up to elytral apex, and often variable inside one population; one of the most typical species character is the presence of long and dense setae tuft at the apex of 3rd antennal joint, which can be completely absent in some northern populations or sometimes in certain southern; sometimes poorly developed setae tuft can be seen at the apex of 4th joint; definite specimens can have more or less contrast grey stripe along humeri; body length in males: 9.5-21.4 mm, in females - 10.5-22.5 mm.

Distribution. From Pyrenean Peninsula to Eastern Siberia; the species is known from Omsk, Tomsk, Novosibirsk and Kemerovo regions, as well as from Altay Region; Kazakhstan (including Zaysan Depression), Central Asia, Caucasus with Transcaucasia, Near East including Iran; in Europe the species does not reach Poland; in East Asia it penetrates to China and Mongolia (Lin & Tavakilian, 2019).

All records for Baykal environs (Cherepanov, 1984; Sama, 2002; Shapovalov, 2012; Danilevky, 2023) were connected with wrong Cherepanov’s identification of Ag. alternans Fischer von Waldheim, 1842.

Biology. Larvae develop in stems of various herbaceous plants: Althaea, Arctium, Cannabis, Carduus, Carthamus, Cichorium, Cirsium, Conium, Daucus, Dictamnus, Dipsacus, Eremurus, Eupatorium, Helianthus, Heracleum, Laserpitium, Latuca, Malva, Melilotus, Onopordum, Pyrethrum, Pastinaca, Sambucus, Sonchus and others; certain populations may clearly exhibit oligophagy or even monophagy; in Europe preference is usually noted for such Asteraceae as Carduus, Cirsium and Onopordum; in Eastern Siberia and Kyrgyzstan they are known only on Malvacea (usually Althaea). According to Danilevsky (2023), in Novorossiysk (Gayduk) and in Crimea (Koktebel) beetles were observed only on Asphodelina. It can be assumed that food specialization demonstrates reproductive isolation of the corresponding populations.

According to Tsherepanov (1984): The entire development cycle is completed within one or two years, because a part of the population pupates after the first winter, and the rest remains for the second winter.

1. Agapanthia (Epoptes) dahlii dahlii (Richter, 1820)

Saperda dahlii Richter, 1822: pl. 12 - “Hungaria”.

New material. 18 males, 7 females, Kazakhstan, Bey-Chogur, Turgay Region, 25.5.-13.6.1916, P. Zikharev & Pania - ZMM; 1 female, Kazakhstan, Akmolinsk Region, lake Usu-Kul, 1912, Maltsev - ZMM; 1 female, Kazakhstan, Akmolinsk Region, Kunduzda River, 29.6.-1.7.1900, Balykleysky leg. - ZIN.

Distribution. North of the species area from Pyrenees to Urals with Sverdlovsk and Tyumen regions; North Caucasus, Crimea, Georgia (part.), North and Central Kazakhstan (part.).

2. Agapanthia (Epoptes) dahlii kuleshovi Danilevsky, 2018

Agapanthia (Epoptes) dahli kuleshovi Danilevsky, 2018: 179 - “Russia, Tomskaya Obl., Belousovo env., 56°18’13"N, 85°11’53"E”, “Tomskaya Obl., Kozhevnikovo Distr., Osinovka (55°57'23"N, 83°29'18"E) env.”; Danilevsky, 2020: 302 - Western Siberia (Tomsk Region); 2023: 583.

New material. 2 males, 1 female, Russia, West Siberia, Kurgan Region, 120 km W Kurgan, 150 m, 4.7.2002, M. Danilevsky - MD; 1 female, 50 km W Kurgan, 24.6.2003, A.A. Safronov - MD; 4 males, 1 female, Omsk Region, Bolsheukovsky District, Bolshiye Uki, 29.5.-9.7.2007, V. Teploukhov - VU; 2 males, 1 female, Bolsheukovsky District, Belogrivka environs, 18.6.2016, V.Н. Teplov - VU.

Distribution. West Siberia of Russia, Kurgan and Tomsk Region.

3. Agapanthia (Epoptes) dahlii efimovi Danilevsky, 2021

Agapanthia dahli, Efimov, 2001: 69 - Kemerovo Region.

Agapanthia (Epoptes) dahli efimovi Danilevsky, 2021: 450 - “Russia, Kemerovo Reg., Prokopyevsk Distr., Karakan Mt. Ridge., 6-7 km from Tykhta”, “Prokopyevsk Distr., 7 km NE Oktyabrsky Lug, 54°17'N, 86°55'E”, “Belovo Distr., NW of Karakan Mt Ridge”, “Kemerovo Distr., Staraya Balakhonka, 55°31'44.1"N, 85°53'23.8"E”, “Krapivinsk Distr., 8 km SSW Saltymakovo”, “Chebulinsk Distr., 9 km S Chumay, mouth of Kozhukh River, 55°39.5'N, 87°49.5'E”, “Chebulinsk Distr., Shestakovo, 55°52'59.8"N, 87°59'8.6"E”, “Chebulinsk Distr., Shestakovo”, “Krapivinsk Dist., 8 km SSW Saltymakovo, 54°45'46"N, 87°1'27"E”; 2023: 583.

New material. 1 male, Kemerovo Region, Kemerovo, Sosnovy Bor, 3.6.2006, D. Sidorov - DE; 1 female, Kemerovo Region, Kemerovo, Kuzbass Botanical Garden, Sukhovskoe Lake, 16-20.6.2003, A. Azimov, E. Maksimenko - DE; 1 female, Kemerovo Region, Chebulinsk District, mouth of river Kozhukh and Kiya, VII.2016, S.Luzyanin - DE; 1 female, Kemerovo Region, Yashkino District, Yurts-Konstantinovs, 56.0659°N 84.9863°E, 2.7.2023, D. Sidorov - DE; 1 male, Russia, Republic of Khakassia, Mayna, 2.7.1970, M. Milov - MD; 3 males, 2 females, Russia, Altay, Gorno-Altaysk (north), 850 m, 20.6.1989, S. Saluk - MD; 1 male, 1 female, Russia, Novosibirsk Region, Kochenevo District, Novomikhaylovka, 8.7.1987, V. Grachev - MD; 1 female, Novosibirsk Region, Maslyanino District, Bubenchikovo, 1954, Lurye - ZMM; 1 female, Novosibirsk Region, Kargat District, Rovenskiy, 6.7.1967, V. Kuznezov - ML; 1 male, 1 female, Siberia, Minussinsk, 22.6.-30.6.1908 - ZMM.

Distribution. West Siberia of Russia (Novosibirsk Region, Altay Republic, Kemerovo Region, Khakassia Republic).

4. Agapanthia (Epoptes) dahlii chemalensis ssp. n.

Fig 1

Type locality. Russia, Republic of Altay, Chemal.

Only one female available; body black with numerous erect black setae; moderately wide; head with dense yellow pubescence, condensed along frons and between antennal bases, rather pale in

Fig 1. Agapanthia (Epoptes) dahlii chemalensis ssp. n.: Holotype, female, Russia, Republic of Altay, Chemal, 20.6.1988, Е. Matveev.

front of eyes; genae about as long as lower eye lobes, covered with yellow pubescence; eyes a little convex, about flat, with deep emargination; the distance between upper eye lobes about equal to the width of 1st antennal joint; frons elongate; antennae relatively thick, protruding beyond elytral apices with 4 joints; 1st and 2nd joints black, other joints red basally and black distally; 3rd joint black for about ¼ of its length and here with distinct setae tuft, consisting of several dense short setae; 4th antennal joint with less developed setae tuft; other antennal joints with a few short erect setae apically; prothorax strongly widened posteriorly, much wider than long; pronotum with wide, dense and bright yellow central stripe; scutellum semicircular, covered with dense yellow pubescence; elytra about 2.9 times longer than wide; with moderately dense yellow pubescence; small glabrous elytral areas nearly indistinct; elytral setae spots more or less distinct; grey humeral elytral stripe absent; elytral apices rounded, erect elytral setae distributed to about elytral middle; ventral body side with very dense and regular yellow pubescence: body length: 14.4 mm; width: 4.5 mm.

Differencial diagnosis. The taxon is close to A. (E.) d. efimovi Danilevsky, 2021, but antennae distinctly thicker, with dense setae tufts rather distinct in 4th joint also; frons with denser pubescence; elytra with sparser paler pubescence.

Material. Holotype, female, Russia, Republic of Altay, Chemal, 20.6.1988, Е. Matveev - MD.

Distribution. Russia, Republic of Altay, Chemal environs.

5. Agapanthia (Epoptes) dahlii calculensis Lazarev, 2013

Agapanthia dahli calculensis Lazarev, 2013b: 128 - North-east Kazakhstan: Sibinka river [49°40’27.56”N, 82°39’13.12”E]; Putintzevo env., 20 km N Zyriyanovsk, Maralikha Mt.; S. Uskaman city; 2014: 278 - North-east Kazakhstan, Sibinka River; Karpiński et al., 2018: 88 - East Kazakhstan Region; Almaty Region; Danilevsky, 2020: 302 - Kazakhstan.

New material. 1 female, Kazakhstan, Marka-Kol District, Chernyaevka, Kalodzhar River, 11.6.1986, V. Shilenkov - MD; 1 male, 1 female, Kazakhstan, Kalbinsky Ridge, Samarka, 600 m, 22.4.2002, M. Danilevsky - MD.

Distribution. North-East Kazakhstan; Sibinka River valley approximately 40 km south of Ust-Kamenogorsk, 49°40'28"N, 82°39'13"E; Mount Maralikha (670 m, 49°50'59"N, 84°22'53"E) in the vicinity of Putintsevo, which is 20 km north of Zyryanovsk; the southern outskirts of Ust-Kamenogorsk (450 m, 49°51'45"N, 82°37'54"E); Berezovka (49°41'N, 83°25'E), 7 km east of Serebryansk; the Larikha Mountains (49°49'6"N, 84°26'31"E), 15 km northeast of Zyryanovsk; Marka-Kol District, Chernyaevka, Kalodzhar River; Kalbinsky Ridge, Samarka, 600 m.

6. Agapanthia (Epoptes) dahlii walteri Reitter, 1898

Figs 2-5

Agapanthia walteri Reitter, 1898: 132 - “Armenien: Erzerum. Kleinasien: Mardin”; Winkler, 1929: 1213, part.; Plavilstshikov, 1932: 194; Zaitzev, 1954: 18 - Borjomi, Armenia, Nakhkrai, M. Asia; Breuning, 1961: 185 - “As. occ. mer.”; Villiers, 1967: 369 - “Asie Mineure, Transcaucasie, Iran”; Abai, 1969: 53 - Iran: Mazandaran, Gorgan, Azarbaidjan, Kermanschahan; Fuchs & Breuning, 1971: 437 - “Anatolie: Yenisehir (Hatay); Zw. Yüksekova u. Semdinli (Hak.); Tunceli”; Kasatkin & Arzanov, 1997: 67 - Chechnya: Petropavlovskaya; Adlbauer, 1988: 289 - “30 km E Bingöl”, “NW Yüksekova, Prov. Hakkari”; Danilevsky, 1993: 39 (= A. salviae, Kazjutshits, 1988); Holecová et al., 2002: 10 - Armenia; Rejzek et al., 2003: 170 - “SE. Turkey: Alannyurt E. Gercüş”; Sama et al., 2005: 130 - Iran, Fars.

Agapanthia dalhi var. erivanica Pic, 1900: 14 - “Arménie: Erivan”.

? Agapanthia dahli var. theryi Pic, 1908: 6 - “Perse: Sultanabad”.

Agapanthia (s. str.) dahli, Pic, 1910: 96 (= erivanica Pic), part.; Aurivillius, 1923: 461 (= cardui, F. = gyllenhali Ganglb. = erivanica Pic = lineatocollis, Muls. = nigricornis F. = theryi Pic), part. - “Mittel- und Südeuropa”, “Armenien”, “Persien”; Plavilstshikov, 1930: 25, 39 (“Syn.: A. lineatocollis Muls. 1863, A. cardui Herbst, 1784, A. nigricornis F. 1792”; “ab. erivanica Pic”), part. - “Europa (von Deutschland bis Spanien, Serbien, Italien u.s.w.), Mittel- und Süd-Russland, Kaukasus, Transkaukasien, Persien, Syrien, Klein-Asien”; 1948: 168, part. - All Armenia; 1968: 121, 147 (= nigricornis F. = erivanica Pic) – (throughout the middle and southern Europe), (In the European part of the USSR... north to Chernigov, Tula, Kazan, Kirov, Perm), (Crimea, Caucasus, all Transcaucasia, northwestern Kazakhstan, Uzbekistan), (Northern Iran, Turkish Armenia, Asia Minor, Syria).

Agapanthia (s. str.) walteri, Pic, 1910: 96 - “Arm., Anat.”; Aurivillius, 1923: 467 - Armenien, Kleinasien; Plavilstshikov, 1930: 24, 39 - “Transkaukasien (Erivan, Daralages u.s.w.), Armenien (Ordubad, Kagyzman, Erzerum), Mardin, Anatolien, Klein-Asien”; 1948: 168, part. - Alagez, Araks Valley, Daralagez, Zangezur; 1968: 121, 146 - Armenia, Nakhichevan Autonomous Soviet Socialist Republic. Central and northeastern Turkey; Lobanov et al.., 1982: 269; Danilevsky & Miroshnikov, 1985: 387, 390.

Agapanthia (s. str.) dahli var. erivanica, Aurivillius, 1923: 461 - “Armenien”.

Agapanthia (s. str.) dahli var. theryi, Aurivillius, 1923: 462 - Persien.

Agapanthia dahli m. erivanica, Breuning, 1961: 185.

Agapanthia dahli m. theryi, Breuning, 1961: 185.

Agapanthia salviae, Kazjutshits, 1988: 584 - Armenian SSR, Khosrovsky reserve, Vedinsky section.

Agapanthia dahli, Winkler, 1929: 1213, part. (including ab. erivanica Pic).

Agapanthia (Agapanthiella) walteri, Pesarini & Sabbadini, 2004: 127.

Agapanthia (Epoptes) walteri, Löbl & Smetana, 2010: 216 - Azerbaijan, Armenia, Georgia, Iran, Turkey; Barimani Varandi et al., 2010: 54 - Iran, Mazandaran; Özdikmen, 2013: 22 - “Turkey: Erzurum, Mardin, Hakkari, Tunceli, Bingöl, Amasya, Karş, Batman”; 2021: 1352; 2024: 1965, 2476 - Black Sea (Amasya, Artvin, Bayburt, Gümüşhan), Central Anatolian Region (Çankırı), Eastern Anatolian Region (Bingöl, Erzurum, Hakkâri, Kars, Muş, Tunceli), Mediterranean Region (Hatay), South-Eastern Anatolian Region (Batman, Mardin, Siirt, Şırnak).

Agapanthia (Epoptes) dahli, Sakenin et al., 2011: 7 - “Iran: Ardabil province: Ardabil”.

Agapanthia dahli walteri, Lazarev, 2013a: 443 - Central Transcaucasia.

Agapanthia (Epoptes) dahli walteri, Danilevsky, 2014: 219; 2020: 302 - Azerbaijan, Armenia, Georgia, Iran, Turkey; 2023: 584 - Armenia, Transcaucasia, Turkey, Iran.

Agapanthia (Epoptes) dahlii walteri, Kasatkin, 2020: 243.

New material. 1 male, Georgia, [Borzhom, Gori District, Tiflis Region, Christof leg. - in Russian] - ZIN; 1 male, 1 female, Iran [Western Persia, Luridtan, Bolkha, 27.5.1914, Nesterov - in Russian] - ZIN; 1 male, 1 female. Iran, Qazvin prov., 4 km W Kouhin, 1500 m, 36°22'N, 49°37'E, 31.5.2015, K. Hodek - MD; 1 male, 1 female, Iran, Kermanshah prov., Shamshir env., 1850 m, 6.6.2018, K. Hodek - KH; 1 female, Iran, W Azerbaijan prov., 2 km W Silvana, 37°26´N, 44°51´E, 1635 m, 30.5.2018, K. Hodek - KH; 1 male, 7 females, Iran, Kordestan prov., 3.5 km E Dizll, 35°22´N, 46°11´E, 1350 m, 25.5.2019, K. Hodek - KH; 14 males, 4 females, Iran, Qazvin prov., 54 km NW of Qazvin, Kouhin env., 36°22'18.0051"N, 49°37'9.0083"E, 1520 m., 28- 29.5.2019, M. Holomčík - MH; 1 male, Iran, Qazvin prov., 4 km W Kouhin, 1500 m, 36°22'18.12"N 49°37'8.82"E, 30.5.2019, D. Navrátil - DN; 5 males, 4 females, Iran, Qazvin prov., 4 km W Kouhin, 1500 m, 36°22'18.12"N, 49°37'8.82"E, 30.5.2019, K. Hodek - KH; 1 male, Iran, Kermanshah prov., Shamshir env., 34°59´14.70“N, 46°25´36.71“E, 1830 m, 26.5.2017, D. Navrátil - DN; 1 male, Iran, Kurdistan, Divandareh-city, Saral, 12.5.2016, Fardin Faizi - MD.

Distribution. Azerbaijan, Armenia, Georgia, Iran, Turkey.

Figs 2-5. Agapanthia (Epoptes) dahlii walteri Reitter, 1898:

2 - 14 males, 4 females, Iran, Qazvin prov., 54 km NW of Qazvin, Kouhin env., 36°22'18.0051"N, 49°37'9.0083"E, 1520 m., 28-29.5.2019, M. Holomčík; 3 - 1 male, 7 females, Iran, Kordestan prov., 3.5 km E Dizll, 35°22´N, 46°11´E, 1350 m, 25.5.2019, K. Hodek; 4 - 1 male, 1 female, Iran, Kermanshah prov., Shamshir env., 1850 m, 6.6.2018, K. Hodek;

5 - 1 female, Iran, W Azerbaijan prov., 2 km W Silvana, 37°26´N, 44°51´E, 1635 m, 30.5.2018, K. Hodek. (Photos by K. Hodek & M. Holomčík).

9. Agapanthia (Epoptes) dahlii ismailovae Lazarev, 2013

Agapanthia dahli ismailovae Lazarev, 2013a: 446 - “North Cucasus, Dagestan, Rutul env.”.

Agapanthia (Epoptes) dahli ismailovae, Danilevsky, 2020: 302 - South of European Russia (Dagestan), Azerbaijan; 2023: 585 - Mountains of Dagestan (the environs of the village of Rutul, 41°32'N, 47°25'E) and northeastern Azerbaijan (the environs of the village of Altyagach, 1030 m, 40°52'32"N, 48°56'23"E).

New material. 1 female, “Aresch / Caucasus / A. Schelkownikow” - ZMM.

Distribution. Russia, Dagestan; Azerbaijan (Altyagach, Aresh).

13. Agapanthia (Epoptes) dahlii salviae Holzschuh, 1975

Figs 10-18

Agapanthia (s. str.) salviae Holzschuh, 1975: 88 - “Elburz, Südseite, 10 km nördlich Karadj”.

Agapanthia (Agapanthiella) salviae, Pesarini & Sabbadini, 2004: 127.

Agapanthia (Epoptes) salviae, Löbl & Smetana, 2010: 216 - Iran; Danilevsky, 2020: 303 - Iran “Elburz, Südseite, 10 km nördlich Karadj”.

Agapanthia (Epoptes) dahli salviae, Lazarev, 2024: 31.

Type locality. Iran, Elburz, south side, 10 km north of Karaj.

The taxonomy position of A. salviae Holzschuh is not clear. The type series as well as many new series collected in the type locality contain several specimens without setae tufts on 3rd antennal joints - a single distinguishing character of A. salviae. All known A. salviae were collected together with A. dahli walteri and all on same food plant (Salvia). Most probably all A. salviae are just a rare morphological form of A. dahli walteri.

New material. 1 male, Iran, Ardabíl prov., Shormineh env., 37°27'N, 48°13'E, 1370 m, 26.5.2018, K. Hodek - KH; 3 males, 1 female, Iran, Ardabíl prov., Askestan env., 37°28'N, 48°39'E, 1750 m, 26.5.2018, K. Hodek - KH; 1 male, Iran, Ardabíl prov., Askestan env., 37°28'20.57"N, 48°39'29.26"E, 1750 m, 27-29.5.2019, D. Navrátil - DN.

Fig. 10. Agapanthia (Epoptes) dahlii salviae Holzschuh, 1975: 3 males, 1 female, Iran, Ardabíl prov., Askestan env., 37°28'N, 48°39'E, 1750 m, 26.5.2018, K. Hodek (Photos by K. Hodek).

Figs 11-15. Agapanthia (Epoptes) dahlii salviae Holzschuh, 1975: male, with same label (11 - view from above, 12 - lateral view, 13 - penis,

14 - parameres, 15 - label) (Photos by K. Hodek).

Figs 16-18. Agapanthia (Epoptes) dahlii salviae Holzschuh, 1975: male, Iran, Ardabíl prov., Shormineh env., 37°27'N, 48°13'E, 1370 m, 26.5.2018, K. Hodek (16 - view from above, 17 - lateral view, 18 - label) (Photos by K. Hodek).

19. Agapanthia (Epoptes) dahlii pustulifera Pic, 1905

Agapanthia pustulifera Pic, 1905: 12 - “Jérusalem”.

Agapanthia (Agapanthiella) pustulifera, Pesarini & Sabbadini, 2004: 127.

Agapanthia (Epoptes) mutinensium Sama & Rapuzzi, 2010b: 179, syn. nov. - “N. Liban, Akkar, Abboudiyeh”; Cocquempot et al., 2016: 99 - Lebanon: “Batloun El Shouf (= Batloun, Mont Liban)”, “Aitat (Aley, Mont Liban)”; Danilevsky, 2020: 303 - Lebanon.

Agapanthia (Epoptes) subsimplicicornis Sama & Rapuzzi, 2010b: 180, syn. nov. - “Liban: Environs de Chtaura”, “Caza Bcharré: Bcharré”, “Bekaa, Ras el Assi, Nahr el Assi, 680 m, N 34°21’06”, E 36°22’36”; Sama et al., 2010a: 34 - Lebanon, ?“Israel - Carmel” (Agapanthia (Epoptes) sp.); Cocquempot et al., 2016: 99 - Lebanon, “Baalbeck, Bekaa”; Danilevsky, 2020: 303 - Lebanon.

Agapanthia (Epoptes) pustulifera, Sama et al., 2010a: 4, 33 - Syria, Jordan, Israel; Sama et al., 2010b: 180 - “Syrie, Jordanie, Liban (?), Israël”; Löbl & Smetana, 2010: 216 - Jordanian, Israel, Lebanon, Syria; Ali & Rapuzzi, 2016: 267 - Syrian Coastal Region; Cocquempot et al., 2016: 99 - “Jafa”, “Batloun (Mont Liban)”; Kasatkin, 2020: 244; Cocquempot et al., 2020: 219 - Syrie: Salkhad (Suweida), Amrit, Hosn, Bilyoun; Danilevsky, 2020: 303 - Jordanian, Israel, Lebanon, Syria.

Agapanthia dahli pustulifera, Lazarev, 2024: 31.

Agapanthia dahli mutinensium, Lazarev, 2024: 31.

Agapanthia dahli subsimplicicornis, Lazarev, 2024: 32.

New material. 1 female, Turkey, Gaziantep, 2 km NWW Fevzipaşa vill., Nurdagi geç., 1100-1140 m, 1.6.2011, A. Napolov & I. Roma - MD.

Distribution. Israel, Syria, Jordan, Lebanon, Turkey (new record).

Note. All three names A. pustulifera, A. mutinensium and A. subsimplicicornis were connected with one variable taxon. Each name was used for specimens from one population. A. pustulifera was observed together with “A. subsimplicicornis” in Bekaa; and with “A. mutinensium” in Batloun. Bisides, according to Cocquempot et al. (2016) A. pustulifera from Bekaa was identified by Sama et al. (2010b) as A. mutinensium. Specimens of Agapanthia mentioned by Cocquempot et al. (2016) were identified by P. Rapuzzi.

21. Agapanthia (Epoptes) dahlii setosa ssp. n.

Figs 6-7

Type locality. Turkey. Isparta, Eğridir environs.

Body black with poor bronze luster; moderately wide; head with dense yellow pubescence, condensed in front of eyes and between antennal bases; genae about as long as lower eye lobes, covered with yellow pubescence; eyes a little convex, about flat, with deep emargination; the distance between upper eye lobes about equal to the width of 1st antennal joint; frons elongate; antennae relatively thin, protruding beyond elytral apices with 4 joints in males or with 3 - in females; 1st and 2nd joints black, other joints red basally and black distally; 3rd joint black for about ¼ of its length with long and dense setae tuft; 4th antennal joint with several long apical setae; other antennal joints with a few short erect setae apically; male prothorax about as wide basally as long; female prothorax slightly wider basally; pronotum with less wide, dense and bright yellow central stripe; scutellum semicircular, covered with dense yellow pubescence; elytra in males about 3 times longer than wide, in females - about 2.9 times; look glabrous, slightly shining; elytral setae spots indistinct; grey humeral elytral stripe absent, but humeral elytral area with shorter and sparser pubescence; elytral apices rounded, short oblique elytral setae distributed to about elytral middle; ventral body side with moderately dense yellow pubescence: body length in males: 12.8-20.3 mm; width: 3.3-5.1 mm; body length in females: 15.3-20.4 mm; width: 4.0-5.6 mm.

Differencial diagnosis. The new subspecies is similar to A. d. lateralis Ganglbauer, 1884 described from “Constantinopel” (Istanbul) because of poor development of elytral pubescence, but body of A. d. lateralis is much wider with usually distinct grey lateral elytral stripe. It strongly differs from A. simplicicornis Reiter, 1898 (described from Mardin) by presence of antennal setae tufts and light 3rd antennal joint; 3rd joint of anterior tarsus is not elongated. A photo of the lectotype of A. simplicicornis Reiter, 1898 similar to A. boeberi (Fischer von Waldheim, 1806) was published by (Kasatkin, 2020).

Figs 6-7. Agapanthia (Epoptes) dahlii setosa ssp. n.: 6 - Holotype, male, Turkey, Isparta, Eğridir, 6.6.1986, S. Kadlec; 7 - Paratype, female, Antalya, Konakli, 25-29.5.2003, K. Vakson.

Material. Holotype, male, [“A. lateralis, det. S. Kadlec”] Turkey, Isparta, Eğridir, 6.6.1986, S. Kadlec - ML; 28 paratypes; male, female [“A. lateralis, det. S. Kadlec”] with same label - ML; 2 males, 1 female, Turkey, Antalya, WSW Kemer, 500 m, 4.5.2010, A. Vlasenko - ML, SM; 7 males, 1 female, Turkey, Antalya, Kemer district, Beklibi env., 13-21.5.2008, 36°44'N, 30°33'E, 15-26.5.2010, A.A. Safronov & D.A. Safronov - MD; 1 male, 1 female, “Asia Minor, Lik. Taurus”, 5.1861, Dr. Schurmann - MD; 1 male, Turkey, Isparta, W Kizilkaya, 25.4.1996, S. Kadlec - MD; 1 male, 3 female, Turkey, Antalya, Seki near Manavgat, 21.5.1996, W. Grosser - MD; 1 male, 1 female, Antalya, Konakli, 25-29.5.2003, K. Vakson - MD; 1 female, Turkey, Içel, NW Erdemli, Aydinlar, 28.5.2001, P. Bialooki - MD; 1 male, 2 females, Turkey, Adana, Hasanbeyli env. pass, 37°07'N, 36°34', 19-25.05.2001, P. Bialooki - MD; 1 female, Hasanbeyli env., N Amanus Mts. 21.5.2001, P. Bialooki - MD; 1 female, Turkey, Buglan geçidi, NW Mus, 17.6.2003, P. Bialooki - MD.

Distribution. Sothern Turkey in several provinces: Antalya, Isparta, Içel, Adana and Mus.

22. Agapanthia (Epoptes) dahlii grossicornis ssp. n.

Figs 8-9

Type locality. Turkey, Çorum.

Body black with poor bronze luster; moderately wide; head with dense yellow pubescence, condensed in front of eyes and between antennal bases; genae about as long as lower eye lobes, covered with yellow pubescence; eyes a little convex, about flat, with deep emargination; the distance between upper eye lobes about equal to the width of 1st antennal joint; frons elongate; antennae distinctly thicker than in A. d. setosa, protruding beyond elytral apices with 4 joints in males or with 3 - in females; 1st and 2nd joints black, other joints red basally and black distally; 3rd joint black for about ¼ of its length; with long and dense setae tuft,; 4th antennal joint without setae tuft, with several long apical setae; other antennal joints with a few short erect setae apically; male prothorax about as wide basally as long; female prothorax slightly wider basally;

Figs 8-9. Agapanthia (Epoptes) dahlii grossicornis ssp. n.: 8 - Holotype, male, Turkey, Çorum, 18.6.1994, N. Auvray; 9 - Paratype, female with same label.

pronotum with less wide, dense and bright yellow central stripe; scutellum semicircular, covered with dense yellow pubescence; elytra in males about 3 times longer than wide, in females - about 2.9 times; look glabrous, slightly shining; elytral setae spots indistinct; grey humeral elytral stripe hardly visible, humeral elytral area with very short and sparse pubescence; elytral apices rounded, short oblique elytral setae distributed to about elytral middle; ventral body side with moderately dense yellow pubescence: body length in males: 15.1-16.5 mm; width: 4.1-4.4 mm; body length in female: 16.5 mm; width: 4.4 mm.

Differencial diagnosis. The new subspecies is similar to A. d. setosa, but differs by rather thick antennae and grey humeral elytral stripe hardly visible; besides it has well developed setae tufts of 3rd antennal joint, doesn’t have setae tufts of 4th antennal joints.

Material. Holotype, male, Turkey, Çorum, 18.6.1994, N. Auvray - ML; 4 paratypes; 2 males, 1 female, with same label - ML, SM; 1 male, Turkey, Amasya, Yenice, 500 m, 21.5.2000, D. Obydov - MD.

Distribution. Northern Turkey, Çorum and Amasya provinces.

27. Agapanthia (Epoptes) dahlii zaysanensis ssp. n.

Figs 19-21

Type locality. East Kazakhstan, Zaysan Lake environs, Zhemeney River.

Body black, elytra without bronze luster; moderately wide; head with dense yellow pubescence, whitish setae are condensed in front of eyes; genae about as long as lower eye lobes, covered with yellow and white pubescence; eyes a little convex, about flat, with deep emargination; the distance between upper eye lobes less than the width of 1st antennal joint; frons a little longer than as wide; antennae rather thin, protruding beyond elytral apices with 2-4 joints; 1st and 2nd joints black, other joints red basally and black distally; 3rd joint black for about ⅓ of its length; with very dense flattened tuft of short black setae; 4th, 5th with similar setae tufts, but considerably reduced; others antennal joints with a few long apical setae; prothorax rather widened basally; pronotum with wide, dense and bright yellow central stripe;

Figs 19-21. Agapanthia (Epoptes) dahlii zaysanensis ssp. n.: 19 - Holotype, male, East Kazakhstan, Zaysan Lake environs, Zhemeney River, 6.6.1910, A. Jacobson; 20 - Holotype label; 21 - Paratype, East Kazakhstan, Zaysan Lake environs, Dzhemeney River, 31.5.1910, A. Jacobson.

scutellum semicircular, covered with dense yellow pubescence; elytra about 2.8 times longer than wide, densely pubescent, often with fused setae patches; grey humeral elytral stripe absent; elytral apices rounded, short oblique elytral setae very short poorly visible along basal third; ventral body side with very dense yellow pubescence: body length in males: 16.0-16.1 mm, width: 3.5-4.0 mm, body length in females: 16.0-18.0 mm; body width: 4.0-4.8 mm.

Differencial diagnosis. The new subspecies is characterized by realtively short antennae, surpassing elytral apex in males with 2-4 joints; setae tufts are distinct on 3rd - 5th antennal joints; seta tufts of 3rd joints very dense, consisting of short setae, protruding all along whole black joint apex.

Material. Holotype, male, East Kazakhstan, Zaysan Lake environs, Zhemeney River, 6.6.1910, A. Jacobson - ZIN; 5 paratypes, 1 male, 2 females East Kazakhstan, Zaysan Lake environs, Dzhemeney River, 31.5.1910, A. Jacobson - ZIN; 2 females with same label - ZMM.

Distribution. East Kazakhstan, Zaysan Lake environs, Zhemeney River.

Etymology. The name of the new taxon is derived from the large lake Zaysan located near the type locality.

28. Agapanthia (Epoptes) dahlii zhidkovi ssp. n.

Figs 22-23

Type locality. Kazakhstan, eastern shore of Alakol lake, 60 km S Makanchi, 408 m, 46°15'24.54"N, 82°12'55.68"E.

Body black, elytra without bronze luster; moderately wide; head with dense yellow pubescence, whitish setae are condensed in front of eyes; genae about as long as lower eye lobes, covered with yellow and white pubescence; eyes a little convex, about flat, with deep emargination; the distance between upper eye lobes less than the width of 1st antennal joint; frons about as long as wide; antennae rather thin, protruding beyond elytral apices with 5 joints in males or with 3 - in females; 1st and 2nd joints black, other joints red basally and black distally; 3rd joint black for about ¼ of its length;

Figs 22-23. Agapanthia (Epoptes) dahlii zhidkovi ssp. n.: 22 - Holotype, male, Kazakhstan, eastern shore of Alakol lake, 60 km S Makanchi, 408 m, 46°15'24.54"N, 82°12'55.68"E, ex pupa, from Eremurus, 9.5.2017, A. Abramov; 23 - Paratypes, female with same label.

with several long apical setae without dense setae tuft; 4th, 5th and others antennal joints with a few long setae; male prothorax about as wide basally as long a little widened basally; female prothorax more widened basally; pronotum with wide, dense and bright yellow central stripe; scutellum semicircular, covered with dense yellow pubescence; elytra in males about 3 times longer than wide, in females - about 2.8 times; densely pubescent, with hardly distinct setae spots; grey humeral elytral stripe absent; elytral apices rounded, short oblique elytral setae very short poorly visible to about apical elytral third; ventral body side with moderately dense yellow pubescence: body length in males: 13.6-14.3 mm; width: 3.4-3.5 mm; body length in females: 14.4-17.4 mm; width: 3.5-3.8 mm.

Differencial diagnosis. The new subspecies is characterized by the absence of setae tufts of 3rd antennal jonts, while elytra densely pubescent with conjugated elytral setae patches; grey lateral elytral stripe absent.

Material. Holotype, male, Kazakhstan, eastern shore of Alakol lake, 60 km S Makanchi, 408 m, 46°15'24.54"N, 82°12'55.68"E, ex pupa, from Eremurus, 9.5.2017, A. Abramov - MD; 9 paratypes; 3 males, 4 females with same label - MD; 1 male, 1 female with same label - ML.

Distribution. East Kazakhstan, eastern shore of Alakol lake, 60 km S Makanchi, 408 m, 46°15'24.54"N, 82°12'55.68"E.

Etymology. The new subspecies is dedicated to Mikhail Borisovich Zhidkov - a friend of Andrey Evgenievich Abramov, who accompanied him in his expedition.

30. Agapanthia (Epoptes) dahlii vishnyakovi ssp. n.

Figs 24-25

Type locality. Uzbekistan, Kashkadarya Region, Zerafshan mountain ridge, Kitab Natural reserve (about 39°10'30"N, 67°18'43"E).

Body black with numerous erect black setae; moderately wide; head with moderately dense yellow pubescence, condensed in front of eyes and between antennal bases; genae about as long as lower eye lobes, covered with yellow pubescence; eyes a little convex, about flat, with deep emargination; the distance between upper eye lobes about equal to the width of 1st antennal joint; frons elongate; antennae relatively thick, protruding beyond elytral apices with 5 joints in males or with 3 - in females; 1st and 2nd joints black, other joints red basally and black distally; 3rd joint black for about ¼ of its length; with distinct setae tuft, consisting of several dense short setae; 4th antennal joint with less developed setae tuft, but 5th joint also has setae tuft, but strongly reduced; other antennal joints with a few short erect setae apically; prothorax moderately widened posteriorly, in males about as long as basal width, in females a little wider posteriorly; pronotum with less wide, dense and bright yellow central stripe; scutellum semicircular, covered with dense yellow pubescence; elytra in males about 3.1 times longer than wide, in females - about 2.6 times; with moderately dense yellow pubescence; many small elytral areas are nearly glabrous; elytral setae spots more or less distinct; grey humeral elytral stripe absent; elytral apices rounded, erect elytral setae distributed to about elytral middle; ventral body side with very dense and regular yellow pubescence: body length in males: 12.4-14.4 mm; width: 2.9-3.5 mm; body length in females: 15.5-16.3 mm; width: 4.3-4.4 mm.

Differencial diagnosis. The new taxon is close to A. d. zaysanensis ssp. n., but differs with dark-grey elytra with scattered pubescence; pronotal setae stripe rather narrow; antennae rather long, surpassing elytral apices in males with 5 joints.

Material. Holotype, male, Uzbekistan, Kashkadarya Region, Zerafshan mountain ridge, Kitab Natural reserve, 6.6.2010 - MD; 6 paratypes; 2 males, 2 females with same label - MD; 1 male with same label - ML; 1 female from same locality, 1.5.2010 - MD.

Distribution. Uzbekistan, Kashkadarya Region, Zerafshan mountain ridge, Kitab Natural reserve (about 39°10'30"N, 67°18'43"E).

Etymology. The new taxon is dedicated to my good friend Alexey Nikolaevich Vishnyakov.

Figs 24-25. Agapanthia (Epoptes) dahlii vishnyakovi ssp. n.: 24 - Holotype, male, Uzbekistan, Kashkadarya Region, Zerafshan mountain ridge, Kitab Natural reserve, 6.6.2010; 25 - female with same label.

31. Agapanthia (Epoptes) dahlii krivosheinae ssp. n.

Figs 26-27

Type locality. Uzbekistan, Syrdarya Region, Yangier.

Two males (one - exceptionally small) and a female available; body black, moderately wide; head with dense yellow pubescence, condensed along frons and between antennal bases, rather pale in front of eyes; genae a little longer than lower eye lobes, covered with yellow pubescence; eyes a little convex, about flat, with deep emargination; the distance between upper eye lobes about equal to the width of 1st antennal joint; frons elongate; antennae relatively thin, protruding beyond elytral apices in big male with 5 joints, in female - with 3 joints; 3rd joint black for about 1/3 of its length and here with long and dens setae tuft; 4th - 5th antennal joints with considerably reduced setae tufts; prothorax strongly widened posteriorly, much wider than long; pronotum with wide, dense and bright yellow central stripe; scutellum semicircular, covered with dense yellow pubescence; elytra in big male about 2.8 times longer than wide, in female - in about 2.5 times; with very dense pubescence, aggregated in numerous small setae spots; small glabrous elytral areas nearly indistinct grey humeral elytral stripes absent; erect elytral setae in big specimens very short and hardly distinct; elytral apices rounded; ventral body side with very less dense yellow pubescence than in other subspecies; body length in males: 10.1-17.9 mm, width: 2.3-4.7 mm; body length in female: 19.2 mm, width: 5.4 mm.

Exceptionally small male has several own characters: antennae just a little longer than body protruding beyond elytral apices with 4 joints; prothorax slightly diverging posteriorly, nearly cylindrical; elytra with distinct long erect, shortly sharpened apically.

Differencial diagnosis. The new taxon is characterized by very long antennae, surpassing elytral apices in males with 5 joints; elytral pubescence very dence, with hardly pronounced setae patches, without grey humeral stripes.

Figs 26-27. Agapanthia (Epoptes) dahlii krivosheinae ssp. n.:

26 - Holotype, male, Uzbekistan, Syrdarya Region, Yangier, 9.5.1980, M. Krivosheina; 27 - Paratypes, female, Uzbekistan, Chatkal Natural Reserve, 22.5.1989, A. Kompatzev.

Material. Holotype, male, Uzbekistan, Syrdarya Region, Yangier, 9.5.1980, M. Krivosheina - ML; 2 paratypes; 1 male, USSR, Uzbekistan, near Syrdarya Station, junction No. 122, 1.5.1931 - ML; 1 female, Uzbekistan, Chatkal Natural Reserve, 22.5.1989, A. Kompatzev - MD.

Distribution. Uzbekistan: Syrdarya Region (Yangier and Syrdarya Station) and Chatkal Natural Reserve.

Etymology. The new taxon is dedicated to doctor of biology sciences Marina Gennadievna Krivosheina, who collected the holotype.

32. Agapanthia (Epoptes) dahlii revadensis ssp. n.

Fig 28

Type locality. Tadzhikistan, Zerafshan valley, Revad environs.

Only one female available; body black with numerous erect black setae; rather wide; head with dense yellow pubescence, condensed in front of eyes and between antennal bases; genae about as long as lower eye lobes, densely covered with yellow pubescence; eyes a little convex, about flat, with deep emargination; the distance between upper eye lobes is about 2 mm; frons slightly elongate; antennae moderately thick, protruding beyond elytral apices with 3 joints; 1st and 2nd joints black, other joints red basally and black distally; 3rd joint black for about ¼ of its length; with distinct setae tuft, consisting of several dense short setae; 4th and 5th antennal joints with a few short erect setae; prothorax moderately widened posteriorly, a little wider than long; pronotum with very wide, dense and bright yellow central stripe; scutellum semicircular, covered with dense and bright yellow pubescence; elytra about 2.8 times longer than wide; with very dense bright yellow continuous pubescence; very small areas with sparse pubescence rather numerous, but elytral spots nearly indistinct; grey humeral elytral stripe absent; elytral apices shortly sharpened; erect elytral setae distributed to about elytral middle; ventral body side with very dense and regular yellow pubescence: body length 20.2 mm; width: 5.2 mm.

Differencial diagnosis. The subspecies is characterized by very bright orange elytral pubescence; elytral setae patches partly fused; setae tuft of 3rd antennal joint pooly developed; 4th and 5th antennal joints with a few short apical erect setae.

Fig. 28. Agapanthia (Epoptes) dahlii revadensis ssp. n.: Holotype, female, Tadzhikistan, Zerafshan valley, Revad environs, 9.6.1994, V. Lukhtanov.

Distribution. A single known locality - Revad (39°23′N 68°11′E) environs in Zerafshan River valley of north-west Tadzhikistan.

Material. Holotype, female, Tadzhikistan, Zerafshan valley, Revad environs, 9.6.1994, V. Lukhtanov - MD.

Etymology. The name of the new taxon comes from the name of the type locality.

A key to the subspecies of Agapanthia (Epoptes) dahlii

(C.F.W. Richter, 1820)

1(36) 4th antennal joint with reduced setae tuft.

2(29) Setae tuft of 3rd antennal joint well developed.

3(18) Elytra with dense and bright pubescence.

4(13) Elytral setae spots very contrast.

5(6) Grey humeral elytral stripe usually well developed; body length: 11.3-20.5 mm. Southwest Azerbaijan, South of Armenia, Iran (East Azerbaijan province).

8. A. d. rubenyani Lazarev, 2013

6(5) Grey humeral elytral stripe absent.

7(8) Prothorax a little widened basally; about 1.2 times wider posteriorly than anteriorly; elytral setae spots usually separated; elytral apices distinctly pointed; pronotal setae stripe wide; body length: 9.5-19.5 mm Turkmenistan, Kopetdag (near Ashgabat, Firyuza, Kara-Kala, Ay-Dere) and border regions of Iran.

14. A. d. transcaspica Pic, 1900

8(7) Prothorax strongly widened basally, about 1.4 times wider posteriorly, than anteriorly

9(12) Elytral pubescence distinctly spotted.

10(11) Setae tufts of 3rd antennal joint well develop; body length: 9.0-22.0 mm. Europe (Albania, Austria, Belgium, Bosnia and Herzegovina, Bulgaria, ?Belarus, Croatia, Russia (Central European Territory and South European Territory), Czech Republic, France (including Corsica and Monaco), Germany, Hungary, Kazakhstan, Macedonia, Moldova, Montenegro, Poland, Romania, Serbia, Slovakia, Slovenia, Spain (including Gibraltar), ?Switzerland, Ukraine), Asia (Georgia, Kazakhstan).

1. A. d. dahlii (Richter, 1820)

11(10) Setae tufts of 3rd antennal joint considerably reduced, represented by several dense long setae; body length: 8.0-16.0 mm. Turkey.

20. A. d. kindermanni Pic, 1905

12(9) Elytral pubescence about uniform with hardly developed setae spots; body length: 14.0-22.0 mm. Kyrgyzstan, Kazakhstan.

24. A. d. alexandris Pic, 1901

13(4) Elytral spots conjugated.

14(17) 3rd antennal joint with numerous black oblique setae; elytral pubescence less uniform with numerous small glabrous spots.

15(16) Elytra with orange pubescence; frons with several black oblique setae; body length: 20.2 mm. Tadjhikistan (Zerafshan valley, Revad environs).

32. A. d. revadensis ssp. n.

16(15) Elytra with yellow pubescence; frons with several black erect setae; body length: 14,9-19,6 mm. West Siberia of Russia: Novosibirsk Region, Altay Republic, Kemerovo Region, Khakassia Republic.

3. A. d. efimovi Danilevsky, 2021

17(14) 3rd antennal joint without black oblique setae; elytral pubescence more or less uniform with several small glabrous spots; body length: 13.0-22.5 mm. West Siberia of Russia: Kurgan Region, Tomsk Region.

2. A. d. kuleshovi Danilevsky, 2018

18(3) Elytra poorly pubescent.

19(20) Antenna relatively thick; body length: 15.1-16.5 mm. Northern Turkey (Çorum Provinces, Amasya Provinces.

22. A. d. grossicornis ssp. n.

20(19) Antenna relatively thin.

21(22) Black parts of 3rd antennal joint much longer, covers about 1/3 of its length; body length: 15.8-18.0 mm. Tajikistan (North-Western Pamir), Afghanistan (Badakhshān province).

33. A. d. ustinovi Danilevsky, 2013

22(21) Black parts of 3rd antennal joint much shorte, covers about 1/4 of its length or shorter.

23(24) Elytral pubescence with strongly scattered patches; body length: 10.0-22.0 mm. Italy (Sicilia).

15. A. d. sicula Ganglbauer, 1884

24(23) Elytral pubescence rather uniform.

25(26) Elytral pubescence nearly indistinct; body length: 10.0-23.0 mm. Albania (Vlorë County, Delvinë District, Kardhiqit Mts., 39°59'36.28"N, 20°06'10.24"E), Bulgaria, Greece, Macedonia.

16. A. d. schurmanni Sama, 1979

26(25) Elytral pubescence very short but distinct.

27(28) Humeral elytral area with shorter and sparser pubescence; body length: 12.8-20.4 mm. Sothern Turkey (Antalya Province, Isparta Province, Içel Province, Adana Province, Mus Province).

21. A. d. setosa ssp. n.

28(27) Humeral elytral area with same pubescence as dorsal elytral side; body length: 10.0-18.0 mm. Southern Kyrgyzstan, Uzbekistan (Iordan, about 39°56'N, 71°45'E, 2300 m), Tajikistan (Indications by Kadyrov (1989) on A. muellneri - Darvoz Range, Zigar, Viskharv).

26. A. d. alaiensis Kratochvíl, 1985

29(2) Setae tuft of 3rd antennal joint reduced (several type specimens of A. d. calculensis including holotype have distinct setae tufts).

30(31) Elytra with grey humeral stripe; body length: 12.0-24.0 mm. Turkey.

18. A. d. lateralis Ganglbauer, 1884

31(30) Elytra without grey humeral stripe, usually with well-developed pubescence.

32(33) Elytral pubescence less developed; prothorax rather widened posteriorly; body length: 11.4-17.9 mm. North-Eastern Kazakhstan.

5. A. d. calculensis Lazarev, 2013

33(32) Elytral pubescence well developed.

34(35) Elytral setae tufts diffused, partly conjgated; body length: 13.6-17.4 mm. East Kazakhstan (eastern shore of Alakol lake, 60 km S Makanchi, 408 m, 46°15'24.54"N, 82°12'55.68"E).

28. A. d. zhidkovi ssp. n.

35(34) Elytral setae tufts rather contrast with glabrous spaces in between.

13. A. d. salviae Holzschuh, 1975

36(1) 4th antennal joint with well-developed setae tuft.

37(46) Elytral pubescence poorly developed; elytra look dark and shining.

38(39) Elytral look totally glabrous, elytral pubescence usually indistinct; body length: 13,5-16.5 mm. East Azerbaijan (Zarat on the Caspian coast; based on a specimen from the collection of C. Holzschuh with his definitions), Eastern Georgia.

7. A. d. nitidipennis Holzschuh, 1984

39(38) Elytra with slightly visible pubescence.

40(41) Antennae dark, basal part of 3rd antennal joint greysh; body length: 14.0-23.2 mm. Northern Iran (Mazandaran Province).

10. A. d. persica Semenov, 1893

41(40) Antenna light, basal part of 3rd antennal joint reddish.

42(43) Apical setae tuft of 4th antennal joint poorly developed with about 9 long setae only; body length: 12.5-19.8 mm. Kyrgyzstan, Uzbekistan, ?China (Xinjiang).

25. A. d. muellneri Reitter, 1898

43(42) Apical setae tuft of 4th antennal joint well with numerous long setae.

44(45) Elytral punctation very dense, many dots conjugated; elytra nearly glabrous, elytral pubescence often indistinct; body length: 13.0-19.0 mm. Jordan, Israel, Lebanon, Syria.

19. A. d. pustulifera Pic, 1905

45(44) Elytral punctation much sparser, dots never conjugated; elytra with distinct pubescence; body length: 14.0-21.0 mm. Northern Iran (Golestan Province).

12. A. d. golestanica Lazarev, Plewa & Jaworski, 2016

46(37) Elytral pubescence well developed; elytra look yellow.

47(58) Prothorax a little widened basally; about 1.2 times wider posteriorly than anteriorly, or sometimes about equally in wide.

48(51) Antennae short, male antennae protruding beyond elytral apex with 4 joints or about one third of elytral length.

49(50) Elytra yellow with very dense pubescence; pronotal setae stripe very wide; body length: 16.0-18.0 mm. East Kazakhstan (Zaysan Lake environs, Zhemeney River).

27. A. d. zaysanensis ssp. n.

50(49) Elytra dark-grey, with scattered pubescence; pronotal setae stripe rather narrow; body length: 12.4-16.3 mm. Uzbekistan (Kashkadarya Region, Zerafshan mountain ridge, Kitab Natural reserve (about 39°10'30"N, 67°18'43"E)).

30. A. d. vishnyakovi ssp. n.

51(48) Male antennae very long, protruding beyond elytral apex with 5 joints about half of elytral length.

52(53) Pronotal setae stripe rather narrow; body length: 14.0-19.7 mm. Kazakhstan (Southern environs of Lake Sorbulak, 43°33'57.65"N, 76°36'24.93"E; Ili River Valley (Dobun pier)).

23. A. d. iliensis Danilevsky, 2018

53(52) Pronotal setae stripe rather wide.

54(55) Elytral spots concentrated in transverse rows; body length: 13.1-17.2 mm. Russia (Dagestan), North-Eastern Azerbaijan (Altıağac).

9. A. d. ismailovae Lazarev, 2013

55(54) Elytral spots are randomly located.

56(57) Grey humeral elytral stripe rather distinct; body length: 17.0-17.7 mm. Azerbaijan (Jalal-Abad District), ?Iran.

11. A. d. lenkorana Lazarev, Plewa & Jaworski, 2016

57(56) Grey humeral elytral stripe absent; body length: 10.1-19,2 mm. Uzbekistan (Syrdarya Region (Yangier and Syrdarya Station) and Chatkal Natural Reserve).

31. A. d. krivosheinae ssp. n.

58(47) Prothorax strongly widened basally, about 1.4 times wider posteriorly, than anteriorly

59(60) Elytra strongly spotted with very distinct contrast bright yellow setae spots; body length: 10.5-20.0 mm. Azerbaijan, Armenia, Georgia, Iran, Turkey.

6. A. d. walteri Reitter, 1898

60(59) Elytral setae spots less distinct, does not look spotted.

61(62) Elytral spots more or less separated; body length: 10.0-22.0 mm. France (Corse), Italy.

17. A. d. malmerendii Sama, 1981

62(61) Most of elytral spots conjugated.

63(64) Antennal setae tuft smaller and shorter; body length: 14.4 mm. Russia (Republic of Altay, Chemal environs).

4. A. d. chemalensis ssp. n.

64(63) Antennal setae tuft long and large; body length: 15.3-17.6 mm. Kazakhstan (Lepsy River), ?China (Xinjiang).

29. A. d. lepsyensis Danilevsky, 2018

Map. 1. Typical localities of Agapanthia dahlii (Richter, 1821) subspecies.

Map 1. Typical localities of Agapanthia dahlii (Richter, 1821) subspecies.

1. A. d. dahlii (Richter, 1820): “Hungaria”; 2. A. d. kuleshovi Danilevsky, 2018: Russia, Tomsk Region, Belousovo environs, 56°18’13"N, 85°11’53"E; 3. A. d. efimovi Danilevsky, 2021: Russia, Kemerovo Region, Prokopyevsk District, Karakan Mt. Ridge., 6-7 km from Tykhta; 4. A. d. chemalensis ssp. n.: Russia, Republic of Altay, Chemal; 5. A. d. calculensis Lazarev, 2013: North-east Kazakhstan, the valley of the Sibinka River about 40 km south of Ust-Kamenogorsk, 49°40'27.56"N, 82°39'13.12"E; 6. A. d. walteri Reitter, 1898: Turkey, Erzurum; 7. A. d. nitidipennis Holzschuh, 1984: Georgia, environs of the Jvari monastery (41°50′19″N, 44°44′02″E) north of Tbilisi; 8. A. d. rubenyani Lazarev, 2013: South Armenia, Megri District, mountains above Shvanidzor, 39°13'N, 46°22'44''E, 1600 m; 9. A. d. ismailovae Lazarev, 2013: Russia, North Cucasus, Dagestan, Rutul environs (41°32'N, 47°25'E); 10. A. d. persica Semenov, 1893: Iran, Eastern and Central Mazanderan province; 11. A. d. lenkorana Lazarev, Plewa & Jaworski, 2016: Azerbaijan, Jalal-Abad District, Andreevka environs (Karazenjir, 39°16'N, 48°30'E, 10 m); 12. A. d. golestanica Lazarev, Plewa & Jaworski, 2016: Iran, Golestan province, 60 km east Minudasht, 460 m, 37°21'36"N 55°55'48"E; 13. A. d. salviae Holzschuh, 1975: Iran, Elburz, south side, 10 km north of Karaj; 14. A. d. transcaspica Pic, 1900: Turkmenistan, Ashgabat; 15. A. d. sicula Ganglbauer, 1884: Italy, Sicilia; 16. A. d. schurmanni Sama, 1979: Greece, Kastoria; 17. A. d. malmerendii Sama, 1981: Italia, Romagna, Portico di Romagna [44°1'N, 11°46'E]; 18. A. d. lateralis Ganglbauer, 1884: Turkey, Istanbul Province, Istanbul (Constantinopel); 19. A. d. pustulifera Pic, 1905: Israel, Jerusalem; 20. A. d. kindermanni Pic, 1905: Southern Turkey; 21. A. d. setosa ssp. n.: Turkey. Isparta, Eğridir environs; 22. A. d. grossicornis ssp. n.: Turkey, Çorum; 23. A. d. iliensis Danilevsky, 2018: Kazakhstan, Almaty Region, southern environs of Lake Sorbulak (43°33'57.65"N, 76°36'24.93"E), 670 m; 24. A. d. alexandris Pic, 1901: Kyrgyzstan, Kyrgyz Ridge (formerly Alexander Ridge); the type most likely comes from the western (Kazakh) part of the ridge; 25. A. d. muellneri Reitter, 1898: Uzbekistan, Tashkent; 26. A. d. alaiensis Kratochvíl, 1985: Southern Kyrgyzstan at the southern border of the Fergana Valley, Kadamjai (40°7'44"N, 71°43'26"E); 27. A. d. zaysanensis ssp. n.: East Kazakhstan, Zaysan Lake environs, Zhemeney River; 28. A. d. zhidkovi ssp. n.: Kazakhstan, eastern shore of Alakol lake, 60 km S Makanchi, 408 m, 46°15'24.54"N, 82°12'55.68"E; 29. A. d. lepsyensis Danilevsky, 2018: Kazakhstan, Lepsy river, 7 km northeast Koilyk (formerly Antonovka), 45°41'36.22"N, 80°17'58.94"E; 30. A. d. vishnyakovi ssp. n.: Uzbekistan, Kashkadarya Region, Zerafshan mountain ridge, Kitab Natural reserve (about 39°10'30"N, 67°18'43"E); 31. A. d. krivosheinae ssp. n.: Uzbekistan, Syrdarya Region, Yangier; 32. A. d. revadensis ssp. n.: Tadzhikistan, Zerafshan valley, Revad environs; 33. A. d. ustinovi Danilevsky, 2013: Tadzhikistan, Pamir, Poshkharv environs [38°24'1"N, 71°9'18"E].

Acknowledgements. I am very grateful for all my friends and colleagues who supplied me with necessary data and materials: Andrey Abramov (Krasnodar Region, Russia), Mikhail Danilevsky (A.N. Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Moscow, Russia), Russia), Dmitry Efimov (Kemerovo, Russia), Aleksey Gusakov (Zoological Museum of Moscow State University, Karel Hodek (Brno, Czech Republic), Michal Holomčík (Lužice, Czech Republic), Sergey Murzin (Moscow, Russia), David Navrátil (Litomyšl, Czech Republic), Gérard Tavakilian (Muséum National d'Histoire Naturelle, Paris, France), Vadim Ustinov (Moscow, Russia). Rostislav Filimonov (Saint-Petersburg, Russia) and Ivan Chigray (Zoological Institute of the Russian Academy of Sciences, Saint-Petersburg, Russia) were responsible for the arrangement of certain photos. Boris Korotyaev (Zoological Institute of the Russian Academy of Sciences, Saint-Petersburg, Russia) is hearty gratitude for his kind assistance.

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Received: 10.01.2025

Accepted: 07.03.2025